[RSS Feed/News] "+" sign in route matching syntax meaning? :+int vs :int what's the difference?

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I see some of XenForo core route formats has this patten
comments/:int<profile_post_comment_id>/

and some of them has this + sign,
:+int<thread_id,title>/post-:+int_p<post_id>

What's the difference?
And where should I look if have to find how these syntax parsing so I could understand?

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